BSc in Biology, University of Patras, 1989; PhD, University of Crete, 1994; Research Fellow, University of Verona, 1991; Research Fellow, University of Parma, 1996-1999; Research Fellow, University of Crete, 1999-2005; Researcher, Institute of Applied and Computational Mathematics, FORTH, 2002- ; Assistant Professor of Physiology, University of Crete, 2005-2017; Associate Professor of Physiology, University of Crete, 2017-2021; Professor of Physiology, University of Crete, 2021.